Albion have handed Bruno a new deal but let four senior stars go - including skipper Gordon Greer.

Club captain Greer, Bobby Zamora, Inigo Calderon and Andrew Crofts have all been released by the Seagulls.

Goalkeepers Niki Maenpaa and Casper Ankergren have been offered new contracts but back-up left-back Adam Chicksen is being let go.

The announcement marks the end of an era as a quartet of senior players whose Albion careers spanned both Withdean and the Amex leave the club.

Of the four, Greer was arguably the player whose future seemed to be most in the balance.

He returned to the side with three solid performances at the end of the season and has also played for Scotland in recent months.

The Scot made 233 starts for Albion, plus one game off the bench, after signing in the summer of 2010 and has been club captain throughout his time with the Seagulls.

Inigo Calderon took his appearance tally to 232 this season, including 202 starts, and scored an impressive 19 goals, mainly from right-back.

His six-and-a-half years will also be remembered for work he did off the pitch with Albion In The Community.

Zamora and Crofts both leave the club for a second time. Zamora’s one-year swansong to his Albion career saw him score seven goals in 26 appearances, of which only ten were starts, before being ruled out by a hip problem.

He scored 90 goals, spanning almost 16 years, in 162 outings for the club in total.

Crofts was Greer’s predecessor as team captain under Gus Poyet and returned to the club four years ago after a stint at Norwich City. He scored 13 goals for the Seagulls in total.

Chicksen’s chances have been more few and far between since signing a three-year deal on arrival from MK Dons in 2013. He played 16 times for the club.